Okay--Situation may be different from when the question was first posed.
You are saying that the plane cannot move forward.
We need to establish some points here or we are stuck in a question of relativity.
The plane cannot move forward relative to what?
Let's build a conveyer belt covering the whole east-west runway. Paint two big dots, a green one and a red one on the conveyer belt right in front of the control tower. Place an airplane on the green dot and a car on the red dot.
The control tower--which is not on the runway or conveyer belt-- stays put. We call that point directly in front of the control tower POINT ZERO.
Now the car and the airplane start to run. Simultaneously the airplane's engine and the cars engine start to drive the respective vehicles west at what should be 200 miles per hour (relative to POINT ZERO). The conveyer bels is dong nothing. Both vehicles move forward (west) at 200 miles per hour relative to POINT ZERO. They leave their respective dots (which are parked at ground zero because the conveyer is not running) behind at that rate.
Now, let's run it again with the conveyer belt. We start the car and the airplane off again to the west at 200 miles per hour. But the COnveyer belt starts to run east at precisely the same time at 200 miles per hour (or 200 miles per hour to the east). SO, the car engine and the airplane each speed up to 300 miles per hour worth of output west. But exactly simultaneously, the conveyer belt speeds up also to 300 miles per hour east.
Is this the situation you are describing?
In this situation, The dots on the conveyer belt are next to each other heading 300 miles per hour east in relation to POINT ZERO. The car is going three hundred miles per hour west in relation to its red dot, and zero miles per hour in relation to the control tower at POINT ZERO. The airplane is going three hundred miles per hour west in relation to the automobile and the control tower at point zero, and six hundred miles an hour west in relation to its green dot.
Now, let us posit another example--the one that you were imagining it appears: Let us say the the conveyer belt is now going to head east at exactly whatever rate the plane is travelling west. If the plane speeds up to a million miles an hour west, the conveyer belt with its little dot will be heading a million miles an hour east.
Unless the conveyer belt itself is attached to the airplane, and vice-versa, then it does not matter how fast the conveyer belt is going. Planes are not attached to conveyer belts.
It is as if you are asking the question: "If I had a perpetual motion machine, could it stay in motion forever." Well, by definition, yes. But, of course, there is no such thing as a perpetual motion machine.
(Note, all of this gedanken experiment assumes efficient wheels and engines, and that there can actually be matching accelerations, and so forth.)
Now, let us posit more than just a conveyer belt. Your conveyer "system" includes the air mass around the airplane. Now, you race the airplane west at exactly 600 miles an hour, but (relative to some imagined external reference point) it appears to be staying perfectly still because everything else out there is moving exactly east at exactly the same 600 miles per hour. That everything includes an airmass that must move east at 600 miles per hour, and if the plane is staying fixed in space, with a six hundred miles per hour headwind, it will fly. So, the plane flies.
Fact is, if the air of sufficient density is flowing with sufficient speed over properly designed wings, the airplane will lift. Whether that airflow comes from the plane moving through the air or the air moving past the wing (after for instance, being forced by a fan in a wind tunnel), the wing will lift the plane.
Alex